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Upeneus tragula Richardson, 1846

Robert K. Whitton Sunday, July 17, 2016 USS Liberty []

Robert K. Whitton Friday, April 21, 2006 Tk2 []

Robert K. Whitton Monday, April 17, 2006 Torpedo Alley []
Richardson 1846 described tragula on page 220 in the 1846 journal article, "Report on the ichthyology of the seas of China and Japan", originally placing the species in the genus Upeneus Cuvier, 1829.

Other Authors using this species concept:

• In Eschmeyer, William N. 2004 Electronic Source The Catalog of Fishes Online as Upeneus tragula
• Page 220 In Richardson, John. 1846 Journal Article Report on the ichthyology of the seas of China and Japan 187-320 as Upeneus tragula
• Page 161,77A,299 In Myers, Robert F. 1999 Book Micronesian reef fishes: a comprehensive guide to the coral reef fishes of Micronesia i-vi, 1-330, pls. 1-192 as Upeneus tragula

Taxonomy Kingdom Animalia  Superphylum Deuterostomia   Phylum Chordata    Subphylum Craniata     Superclass Gnathostomata      Class Actinopterygii       Order Perciformes        Suborder Percoidei         Family Mullidae          Genus Upeneus           Species tragula

Locations

Evidence 16 VideoRecordings 2 StillImages 1 Reference 2-106m

Associated Organisms

Taxa	Identified As	Location	Evidence Type	Event Date	Observers/Collectors	Determiners	Min Depth	Max Depth
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Torpedo Alley	Video Recording	04/17/2006	Robert K. Whitton	Robert K. Whitton	14m	14m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Torpedo Alley	Video Recording	04/17/2006	Robert K. Whitton	Robert K. Whitton	14m	14m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Tk2	Video Recording	04/21/2006	Robert K. Whitton	Robert K. Whitton	12m	12m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Tk2	Video Recording	04/21/2006	Robert K. Whitton	Robert K. Whitton	14m	14m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Tangeng Pekat	Video Recording	04/24/2006	Robert K. Whitton	Robert K. Whitton	2m	2m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Anilao	Video Recording	01/31/2010	John L. Earle	John L. Earle	9m	9m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Basura	Video Recording	03/28/2009	John L. Earle	John L. Earle	6m	6m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Basura	Video Recording	03/24/2009	John L. Earle	John L. Earle	8m	8m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Raja Ampat	Video Recording	10/09/2005	John L. Earle	John L. Earle	18m	18m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Palau Islands	Reference	01/01/1999	Myers 1999 Page(s) 299	Micronesian reef fishes: a comprehensive guide to the coral reef fishes of Micronesia Coral Graphics,Barrigada, Guam. Third edition, revised and expanded Edition: i-vi, 1-330, pls. 1-192.	m	m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Tulamben	Video Recording	08/23/2015	John L. Earle	John L. Earle	9m	9m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Tulamben	Video Recording	07/22/2016	Robert K. Whitton	Robert K. Whitton	8m	8m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Tulamben	Video Recording	07/22/2016	John L. Earle	John L. Earle	7m	7m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	USS Liberty	Video Recording	07/18/2016	Robert K. Whitton	Robert K. Whitton	18m	18m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Dalig Point ( Dive and Trek Reef )	Video Recording	12/12/2013	Robert K. Whitton	Richard L. Pyle	103m	106m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	South Twin Rocks	Video Recording	01/26/2010	Robert K. Whitton	Robert K. Whitton	15m	15m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Bluri Village	Video Recording	05/08/2012	Robert K. Whitton	Robert K. Whitton	7m	7m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Anilao Pier	Still Image	02/14/2026	Robert K. Whitton	Robert K. Whitton	9m	9m
Upeneus tragula Richardson, 1846	Upeneus tragula Richardson, 1846	Cathedral	Still Image	02/14/2026	Robert K. Whitton	Robert K. Whitton	16m	16m

Names linked to the original usage
Upeneus tragula Richardson, 1846 Species
Upeneoides kuiskuiana Döderlein in Steindachner & Döderlein, 1883 Species
Upeneus kuiskuiana (Döderlein in Steindachner & Döderlein, 1883) Species
Upeneus niebuhri Guézé, 1976 Species
Upeneus oligospilus Lachner, 1954 Species
Upeneoides variegatus Bleeker, 1849 Species
Upeneus variegatus (Bleeker, 1849) Species